The oceanic Cocos Plate subducting beneath Costa Rica has a complex plate tectonic history resulting in segmentation. New lines of magnetic data clearly define tectonic boundaries which separate lithosphere formed at the East Pacific Rise from lithosphere formed at the Cocos-Nazca spreading center. They also define two early phase Cocos-Nazca spreading regimes and a major propagator. In addition to these sharply defined tectonic boundaries are overprinted boundaries from volcanism during passage of Cocos Plate over the Galapagos hot spot. The subducted segment boundaries correspond with distinct changes in upper plate tectonic structure and features of the subducted slab. Newly identified seafloor-spreading anomalies show oceanic lithosphere formed during initial breakup of the Farallon Plate at 22.7 Ma and opening of the Cocos-Nazca spreading center. A revised regional compilation of magnetic anomalies allows refinement of plate tectonic models for the early history of the Cocos-Nazca spreading center. At 19.5 Ma a major ridge jump reshaped its geometry, and after ∼14.5 Ma multiple southward ridge jumps led to a highly asymmetric accretion of lithosphere. A suspected cause of ridge jumps is an interaction of the Cocos-Nazca spreading center with the Galapagos hot spot.